Saturniidae of Borneo
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 compiled by Wolfgang A. Nässig

A treatise on the Saturniidae (and Brahmaeidae, Eupterotidae, Sphingidae etc.) of the island of Borneo (East Malaysia and Indonesia) was published by J.D. Holloway in 1987:
Jeremy D. Holloway (1987): The Moths of Borneo, Part 3: Superfamily Bombycoidea: families Lasiocampidae, Eupterotidae, Bombycidae, Brahmaeidae, Saturniidae, Sphingidae.

Kuala Lumpur (Southdene), 199 pp., 163 genitalia figs., 20 col. pls.

Availability & prices: see here.

The Moths of Borneo Part 3

The following species of Saturniidae are known from Borneo:
Systematic arrangement according to the order used in the more recent Sumatran, which shares a lot of species with Borneo, and Philippine Saturniidae faunas. Additions or changes in comparison to Holloway’s list of 1987 are listed as footnotes following the list. The systematics and nomenclature applied here are therefore not identical with the original treatment by Holloway in 1987! — See also the disclaimer below. — References see at the end.

Holloway (1987) listed 22 species of Saturniidae for the island of Borneo; here we show 23 species based on the present knowledge.

Superfamily Bombycoidea Latreille, 1802
Family Saturniidae Boisduval, [1837] 1834
Subfamily Saturniinae Boisduval, [1837] 1834

Tribe Attacini Blanchard, 1840
Genus Attacus Linnaeus, 1767

Attacus atlas (Linnaeus, 1758)
Genus Archaeoattacus Watson, 1914
Archaeoattacus staudingeri (Rothschild, 1895)
Genus Samia Hübner, [1819] 1816
Samia (insularis) tetrica (Rebel, 1924) (see footnote 1)
Tribe Saturniini Boisduval, [1837] 1834
Genus Actias Leach, 1815
Actias maenas (Doubleday, 1847) (see footnote 2)
Actias selene (Hübner, 1806)  (see footnote 3)
Genus Cricula Walker, 1855
Cricula elaezia Jordan, 1909
Cricula bornea Watson, 1913 (see footnote 4)
Cricula trifenestrata (Helfer, 1837)  (see footnote 5)
Genus Antheraea Hübner, [1819] 1816
Subgenus Antheraeopsis Wood-Mason, 1886
Antheraea (Antheraeopsis) youngi Watson, 1915 (see footnote 6)
Subgenus Antheraea Hübner, [1819] 1816
Antheraea (Antheraea) larissa (Westwood, 1847)
Antheraea (Antheraea) platessa Rothschild, 1903 (nec jana (Stoll, 1782)) (see footnote 7)
Antheraea (Antheraea) gschwandneri Niepelt, 1918 (species-complex?) (see footnote 8)
Antheraea (Antheraea) brunei Allen & Holloway, 1986
Antheraea (Antheraea) alleni Holloway, 1987 (see footnote 9)
Antheraea (Antheraea) moultoni Watson, 1927 (see footnote 10)
Antheraea (Antheraea) rosieri (Toxopeus, 1940)
Antheraea (Antheraea) helferi borneensis Moore, 1892 (see footnote 11)
Antheraea (Antheraea) diehli Lemaire, 1979
Antheraea (Antheraea) (pernyi) roylii korintjiana Bouvier, 1928  (see footnote 12)
Antheraea (Antheraea) lampei Nässig & Holloway, 1989 (see footnote 12)
Genus Loepa Moore, 1859 (see footnote 13)
Loepa megacore Jordan, 1911
Loepa sikkima javanica Mell, 1938 (see footnote 14)
Genus Lemaireia Nässig & Holloway, 1987 (see footnote 15)
Lemaireia loepoides (Butler, 1880)


Important Note/Disclaimer: The taxonomic changes proposed here within this web page are not proposed as valid taxonomic acts in accordance with the Code; they are only suggestions to promote the ongoing discussion among the specialists.

References see below!

  1. This species was listed by Holloway as Samia tetrica. The Samia taxa of Sundaland (except the second Peninsular Malaysian species?) all belong to the insularis complex. The interpretation which taxon is to be treated as species or subspecies remains to be assessed; a revision of the whole genus is in preparation by R.S. Peigler and S. Naumann. — Back.
  2. The subspecific status of Bornean (and other Sundanian) Actias maenas populations requires further research; within the Sumatra fauna the Sundaland population was called A. maenas diana Maassen, [1872]. — Back.
  3. The subspecific status of Bornean (and other Sundanian) Actias selene populations requires further research; within the Sumatra fauna the Sundaland population was called A. selene seitzi Kalis, 1934. As was recently shown by U. & L.H. Paukstadt (1999a), the populations of the selene-group from Java and the other parts of Sundaland are not identical. The valid subspecific name for the Bornean population of A. selene may be vandenberghi; but further research remains necessary. — Back.
  4. The status of Cricula bornea is still unclear. — Back.
  5. The subspecific status of the Bornean population of Cricula trifenestrata as well as the differences with C. bornea still require further research. — Back.
  6. Holloway called this species A. assamensis Helfer, 1837. However, the true A. (Ao.) assamensis is restricted to the Himalayan area (and maybe mountains in northern Indochina/South China?) (studies are in preparation, see under the website In the Sumatra treatment of 1996, this taxon youngi Watson, 1915 (applied to the Sumatran population) was interpreted as a subspecies of castanea Jordan, 1910. It was then raised to full species status by Nässig & Treadaway (July 1998, “The Saturniidae (Lepidoptera) of the Philippines”). See also U. Paukstadt et al. (August 1998), who published some additional information on the Sumatran fauna of Antheraea. — Back.
  7. In 1987, when Holloway’s Borneo publication was published, the status and identity of “jana” sensu auctorum was not yet resolved. This was only done by Nässig (1992). Holloway’s “A. jana” in fact is A. platessa. The true A. jana is a much smaller species evidently endemic to Java and Bali. — L.H. & U. Paukstadt (1999) described the larva of the Bornean A. platessa. It is quite similar to all other species of the frithi-subgroup, but shows distinctive dichromatic dorsal scoli on meso- and metathorax in last instar. Regrettably it was not stated whether the dorsal scoli on A8 are separated in last instar (as illustrated for earlier instars) or fused. — Back.
  8. Holloway called this Bornean population A. celebensis Watson, 1915. This name celebensis, however, should be restricted to the Sulawesian population. Calling this Bornean species gschwandneri here in the present web page follows the concept applied in the Sumatran web page. However, there are other concepts, see the footnote in the Sumatran web page. The situation within the species of the frithi-subgroup (how many real biological species are included in this group, and how can we differentiate them?) is still far from being solved, see also the following footnotes. — The species A. ulrichbroschi U. & L.H. Paukstadt, 1999, described from Peninsular Malaysia, may possibly also live on Borneo, in case that it is a distinct species; compare U. & L.H. Paukstadt (1999b). — Back.
  9. See the following footnote. — Back.
  10. A. alleni and A. moultoni are two more taxa which also belong to the frithi-complex. Although these populations appear to be easily recognizeable externally, we still lack a safe confirmation of the species status on a reliable and well-done methodological base. — Back.
  11. The original helferi Moore, 1859 was described from India. The status of the Sundanian populations requires further research. — Back.
  12. When the publication by Holloway was published in 1987, the identity of these two taxa was not yet solved; Holloway dealt with both species under the name of “A. korintjiana Bouvier, 1928” only. In a later paper (Nässig & Holloway 1989) we separated these two taxa. See also the remarks in the footnote of the Sumatran publication. — Back.
  13. This taxon was described first by Moore in 1859, not in his “1858–1859” [1860] publication, see also the remarks about these publications in the footnote of the Sumatran publication. The situation within the pernyi-group still has some problems. — Back.
  14. This taxon was called L. sikkima Moore, 1865 by Holloway. However, there are some differences between the nominotypical northern populations in the Himalaya area and the Sundanian ones. — Back.
  15. This genus was inadvertedly described within the publication by Holloway (1987) in advance of the so-intended publication, because the print of the publication by Nässig & Holloway in Heterocera Sumatrana became delayed into early 1988. — Back.

  • Nässig, W. A. (1992): Antheraea (Antheraea) platessa Rothschild 1903: The correct name for Antheraea jana auctorum, nec Stoll 1782 (Lepidoptera: Saturniidae). — Nachr. entomol. Ver. Apollo, Frankfurt/Main, N.F. 13 (2a): 157–163.
  • Nässig, W.A., & Holloway, J.D. (1988) On the systematic position of “Syntherata” loepoides Butler and its allies (Lep., Saturniidae). — Heterocera Sumatrana 2 (6): 115–127)
  • Nässig, W.A., & Holloway, J.D. (1989) A new species of Antheraea from Sumatra, West Malaysia and Borneo (Lepidoptera, Saturniidae). — Heterocera Sumatrana 2 (7): 185–190.
  • Paukstadt, L.H., & Paukstadt, U. (1999): Die Beschreibung der Präimaginalstadien von Antheraea (Antheraea) platessa W. Rothschild, 1903 von Nord Borneo [sic], sowie Angaben zur Biologie und Ökologie (Lepidoptera: Saturniidae). — Galathea, Ber. Krs. Nürnberger Entomol., Suppl. 6: 13–22.
  • Paukstadt, U., & Paukstadt, L.H. (1999a): Anmerkungen zum taxonomischen Status von Actias selene seitzi Kalis 1934 und Actias selene vandenberghi Roepke 1956 (stat. rev.), sowie zur geographischen Verbreitung dieser Taxa in Südostasien (Lepidoptera: Saturniidae). — Entomologische Zeitschrift 109 (12): 484–491.
  • Paukstadt, U., & Paukstadt, L.H. (1999b) [28. December]: Antheraea (Antheraea) ulrichbroschi n. sp., eine neue Saturniide von West Malaysia [sic] (Lepidoptera: Saturniidae). — Galathea, Ber. Krs. Nürnberger Entomol., Suppl. 6: 90–98.
  • Paukstadt, U., Paukstadt, L.H., & Brosch, U. (1998):Taxonomische Änderungen und Anmerkungen zu den Taxa der Gattung Antheraea Hübner [1819] von Sumatera [sic], Indonesien (Lepidoptera: Saturniidae). — Entomologische Zeitschrift 108 (8): 317–327.

© 1999–2002 by W. A. Nässig — compiled & published 25. October/13. December 1999, last changes 29. December 1999, 18. January 2002
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